Plain English Meaning
The pilot who is in charge of the aircraft. They make the final decisions, carry the responsibility for everything that happens on the flight, and answer for the safety of the people and the aircraft.
Full Definition
The pilot responsible for the operation and safety of an aircraft during flight time. The pilot in command has final authority over the aircraft, is directly responsible for and is the final authority as to the operation of that aircraft, and may deviate from any rule to the extent required to meet an in-flight emergency.
Context Anchor
You will see this term in regulations, flight planning, cockpit decision-making, and accident or incident discussions.
Derivation
From the older military and use of 'in command,' meaning the ultimate authority over a vessel or unit. borrowed the phrase directly: the pilot 'in command' is the one whose word is final, just as a ship's captain is at sea.